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What is an arbuscle?

What is an arbuscle? Here are some definitions.

Noun
  1. (botany) A plant midway in height between a shrub and a tree; a dwarf tree.
  2. (mycology) A branched hypha in some fungi.
  3. (mycology) The site at which a symbiotic fungus attaches to the roots of a plant and exchanges nutrients, etc., with it.
  4. (zoology) A clump of feather-like cilia (hairlike structures).
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