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What is a lacuna?

What is a lacuna? Here are some definitions.

Noun
  1. A small opening; a small pit or depression; a small blank space; a gap or vacancy; a hiatus.
  2. An absent part, especially in a book or other piece of writing, often referring to an ancient manuscript or similar such.
  3. (microscopy) A space visible between cells, allowing free passage of light.
  4. (translation studies) A language gap, which occurs when there is no direct translation in the target language for a lexical term found in the source language
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Examples
Rather, this gap must be seen as a serious lacuna in the surviving texts, a gulf that will critically limit our analysis.
Because of this lacuna, Frickenhaus posited that the text originally described the figure as being by Lysippos.
In this provocative study, Newhauser fills a lacuna in historical scholarship even as he provides insight for the nonhistorian.
Thus, divergent growth apparently prompted offsetting, in order for the coral to maintain the lacuna and occupy the space around it.
Marg's new book is an attempt to fill the lacuna, and comes 25 years after an earlier study titled Homage to Jaipur.
As often bedevils the translator's task there appears to be at least one lacuna in the extant text, but it was relatively painless to decipher.

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