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What is a rune?

What is a rune? Here are some definitions.

Noun
  1. A letter, or character, belonging to the written language of various ancient Germanic peoples, especially the Scandinavians and the Anglo-Saxons.
  2. A Finnish or Scandinavian epic poem, or a division of one, especially a division of the Kalevala.
  3. A letter or mark used as mystical or magic symbol.
  4. A verse or song, especially one with mystical or mysterious overtones; a spell or an incantation.
  5. (obsolete) A roun (secret or mystery).
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If you compare the futhorc given above, you will see that Tolkien has altered some the rune shapes.
There was a Danish king of that name who was famed for his generosity, to whom the rune poem probably refers.
Furthermore, several rune alphabets or futharks have been left behind to puzzle scholars.
If I was as drunk as these bottles say I was, I couldn't have cast a rune, let alone a spell.
Oddly enough, I memorized the Anglo-Saxon rune alphabet in high school with a friend of mine so we could pass notes in geometry class.
Every stone of the edifice was marked with a rune, coursing with magical strength.

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